I can see what you are saying, the top two tuner holes are different distances from the edges. The mids look right as do the ones closest to the nut but the top two are definitely out.
That’s a definite dowel and redrill job unless you want to flick it back to Adam, or forget about it if you can put up with it.
Personally speaking, even the photo of it is giving me a tic, so I’d be cutting the plug for it as we speak. 😵😂